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Cheesebridge is a posh Victorian-era town obsessed with wealth, class, and the stinkiest of fine cheeses. Beneath its charming cobblestone streets dwell the Boxtrolls, an underground cavern-dwelling community of quirky and lovable oddballs who wear recycled cardboard boxes the way turtles wear their shells. The Boxtrolls have raised an orphaned human boy, Eggs (voiced by Isaac Hempstead-Wright), since infancy as one of their dumpster-diving and mechanical junk-collecting own. When the Boxtrolls are targeted by villainous pest exterminator Archibald Snatcher (Academy Award winner Ben Kingsley), who is bent on eradicating them as his ticket to Cheesebridge society, the kindhearted band of tinkerers must turn to their adopted charge and adventurous rich girl Winnie (Elle Fanning) to bridge two worlds amidst the winds of change – and cheese.